Effect of randomization on the robustness effect of gap junctions. Robustness effect was quantified as in Figure 11A. (A) Effect of changing the randomness in the leak currents and cellular diameters (cell randomness) relative to standard (feedback configuration with 20% parallel fiber (PF) synaptic weight). (B) Effect of combined changes to the cell-randomness and the randomness in the spatial position of the cells relative to the standard. (C) Effect of the lack of spatial randomness (red) or of sparser MF inputs onto Golgi cells (GoCs)(black) compared with standard (blue).